The link you followed will bring you to this page:

https://www.antwiki.org/wiki/images/0/01/Ito%2C_F.%2C_Hashim%2C_R._et_al._2021._Colony_composition_in_the_Oriental_ectatommine_ant%2C_Gnamptogenys_menadensis_%2810.20362%40am.013006%29.pdf